I should have been more specific for the sake of clarity:

"Each individual saddle is reversible depending on what gives furthest travel for closest 'best' intonation".

Btw, next time I try to sort out this guitar
I'll probably fit a set of heavier 11's strings
with a higher less comfortable action.
See if that'll tame the over sharp intoned Bass E string.

As a bonus, it'll definitely sound chunkier for power chords & riffs.